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Origin of Species
chapter x. on the imperfection of the geological record   On the lapse of time, as inferred from the rate of denudation and of deposition
Charles Darwin
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       Independently of our not finding fossil remains of such infinitely numerous connecting links, it may be objected that time cannot have sufficed for so great an amount of organic change, all changes having been effected slowly. It is hardly possible for me to recall to the reader who is not a practical geologist, the facts leading the mind feebly to comprehend the lapse of time. He who can read Sir Charles Lyell's grand work on the Principles of Geology, which the future historian will recognise as having produced a revolution in natural science, and yet does not admit how vast have been the past periods of time, may at once close this volume. Not that it suffices to study the Principles of Geology, or to read special treatises by different observers on separate formations, and to mark how each author attempts to give an inadequate idea of the duration of each formation, or even of each stratum. We can best gain some idea of past time by knowing the agencies at work; and learning how deeply the surface of the land has been denuded, and how much sediment has been deposited. As Lyell has well remarked, the extent and thickness of our sedimentary formations are the result and the measure of the denudation which the earth's crust has elsewhere undergone. Therefore a man should examine for himself the great piles of superimposed strata, and watch the rivulets bringing down mud, and the waves wearing away the sea-cliffs, in order to comprehend something about the duration of past time, the monuments of which we see all around us.
       It is good to wander along the coast, when formed of moderately hard rocks, and mark the process of degradation. The tides in most cases reach the cliffs only for a short time twice a day, and the waves eat into them only when they are charged with sand or pebbles; for there is good evidence that pure water effects nothing in wearing away rock. At last the base of the cliff is undermined, huge fragments fall down, and these remaining fixed, have to be worn away atom by atom, until after being reduced in size they can be rolled about by the waves, and then they are more quickly ground into pebbles, sand, or mud. But how often do we see along the bases of retreating cliffs rounded boulders, all thickly clothed by marine productions, showing how little they are abraded and how seldom they are rolled about! Moreover, if we follow for a few miles any line of rocky cliff, which is undergoing degradation, we find that it is only here and there, along a short length or round a promontory, that the cliffs are at the present time suffering. The appearance of the surface and the vegetation show that elsewhere years have elapsed since the waters washed their base.
       We have, however, recently learned from the observations of Ramsay, in the van of many excellent observers--of Jukes, Geikie, Croll and others, that subaerial degradation is a much more important agency than coast-action, or the power of the waves. The whole surface of the land is exposed to the chemical action of the air and of the rainwater, with its dissolved carbonic acid, and in colder countries to frost; the disintegrated matter is carried down even gentle slopes during heavy rain, and to a greater extent than might be supposed, especially in arid districts, by the wind; it is then transported by the streams and rivers, which, when rapid deepen their channels, and triturate the fragments. On a rainy day, even in a gently undulating country, we see the effects of subaerial degradation in the muddy rills which flow down every slope. Messrs. Ramsay and Whitaker have shown, and the observation is a most striking one, that the great lines of escarpment in the Wealden district and those ranging across England, which formerly were looked at as ancient sea-coasts, cannot have been thus formed, for each line is composed of one and the same formation, while our sea-cliffs are everywhere formed by the intersection of various formations. This being the case, we are compelled to admit that the escarpments owe their origin in chief part to the rocks of which they are composed, having resisted subaerial denudation better than the surrounding surface; this surface consequently has been gradually lowered, with the lines of harder rock left projecting. Nothing impresses the mind with the vast duration of time, according to our ideas of time, more forcibly than the conviction thus gained that subaerial agencies, which apparently have so little power, and which seem to work so slowly, have produced great results.
       When thus impressed with the slow rate at which the land is worn away through subaerial and littoral action, it is good, in order to appreciate the past duration of time, to consider, on the one hand, the masses of rock which have been removed over many extensive areas, and on the other hand the thickness of our sedimentary formations. I remember having been much struck when viewing volcanic islands, which have been worn by the waves and pared all round into perpendicular cliffs of one or two thousand feet in height; for the gentle slope of the lava-streams, due to their formerly liquid state, showed at a glance how far the hard, rocky beds had once extended into the open ocean. The same story is told still more plainly by faults--those great cracks along which the strata have been upheaved on one side, or thrown down on the other, to the height or depth of thousands of feet; for since the crust cracked, and it makes no great difference whether the upheaval was sudden, or, as most geologists now believe, was slow and effected by many starts, the surface of the land has been so completely planed down that no trace of these vast dislocations is externally visible. The Craven fault, for instance, extends for upward of thirty miles, and along this line the vertical displacement of the strata varies from 600 to 3,000 feet. Professor Ramsay has published an account of a downthrow in Anglesea of 2,300 feet; and he informs me that he fully believes that there is one in Merionethshire of 12,000 feet; yet in these cases there is nothing on the surface of the land to show such prodigious movements; the pile of rocks on either side of the crack having been smoothly swept away.
       On the other hand, in all parts of the world the piles of sedimentary strata are of wonderful thickness. In the Cordillera, I estimated one mass of conglomerate at ten thousand feet; and although conglomerates have probably been accumulated at a quicker rate than finer sediments, yet from being formed of worn and rounded pebbles, each of which bears the stamp of time, they are good to show how slowly the mass must have been heaped together. Professor Ramsay has given me the maximum thickness, from actual measurement in most cases, of the successive formations in different parts of Great Britain; and this is the result:
       Feet Palaeozoic strata (not including igneous beds).. 57,154 Secondary strata................................ 13,190 Tertiary strata................................. 2,240
       --making altogether 72,584 feet; that is, very nearly thirteen and three-quarters British miles. Some of these formations, which are represented in England by thin beds, are thousands of feet in thickness on the Continent. Moreover, between each successive formation we have, in the opinion of most geologists, blank periods of enormous length. So that the lofty pile of sedimentary rocks in Britain gives but an inadequate idea of the time which has elapsed during their accumulation. The consideration of these various facts impresses the mind almost in the same manner as does the vain endeavour to grapple with the idea of eternity.
       Nevertheless this impression is partly false. Mr. Croll, in an interesting paper, remarks that we do not err "in forming too great a conception of the length of geological periods," but in estimating them by years. When geologists look at large and complicated phenomena, and then at the figures representing several million years, the two produce a totally different effect on the mind, and the figures are at once pronounced too small. In regard to subaerial denudation, Mr. Croll shows, by calculating the known amount of sediment annually brought down by certain rivers, relatively to their areas of drainage, that 1,000 feet of solid rock, as it became gradually disintegrated, would thus be removed from the mean level of the whole area in the course of six million years. This seems an astonishing result, and some considerations lead to the suspicion that it may be too large, but if halved or quartered it is still very surprising. Few of us, however, know what a million really means: Mr. Croll gives the following illustration: Take a narrow strip of paper, eighty-three feet four inches in length, and stretch it along the wall of a large hall; then mark off at one end the tenth of an inch. This tenth of an inch will represent one hundred years, and the entire strip a million years. But let it be borne in mind, in relation to the subject of this work, what a hundred years implies, represented as it is by a measure utterly insignificant in a hall of the above dimensions. Several eminent breeders, during a single lifetime, have so largely modified some of the higher animals, which propagate their kind much more slowly than most of the lower animals, that they have formed what well deserves to be called a new sub-breed. Few men have attended with due care to any one strain for more than half a century, so that a hundred years represents the work of two breeders in succession. It is not to be supposed that species in a state of nature ever change so quickly as domestic animals under the guidance of methodical selection. The comparison would be in every way fairer with the effects which follow from unconscious selection, that is, the preservation of the most useful or beautiful animals, with no intention of modifying the breed; but by this process of unconscious selection, various breeds have been sensibly changed in the course of two or three centuries.
       Species, however, probably change much more slowly, and within the same country only a few change at the same time. This slowness follows from all the inhabitants of the same country being already so well adapted to each other, that new places in the polity of nature do not occur until after long intervals, due to the occurrence of physical changes of some kind, or through the immigration of new forms. Moreover, variations or individual differences of the right nature, by which some of the inhabitants might be better fitted to their new places under the altered circumstance, would not always occur at once. Unfortunately we have no means of determining, according to the standard of years, how long a period it takes to modify a species; but to the subject of time we must return.
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Introduction
chapter i. variation under domestication
   Causes of Variability
   Effects of Habit and the use or disuse of Parts; Correlated Variation; Inheritance
   Character of Domestic Varieties; Difficulty of distinguishing between Varieties and Species; Origin of Domestic Varieties from one or more Species
   Breeds of the Domestic Pigeon, Their Differences and Origin
   Principles of Selection, anciently followed, their Effects
   Unconscious Selection
   Circumstances favourable to Man's power of Selection
chapter ii. variation under nature
   Variability
   Individual Differences
   Doubtful species
   Wide ranging, much diffused, and common species, vary most
   Species of the larger genera in each country vary more frequently than the species of the smaller genera
   Many of the species of the larger genera resemble varieties in being very closely, but unequally, related to each other, and in having restricted ranges.
   Summary
chapter iii. struggle for existence
   Its bearing on natural selection
   The term, Struggle for Existence, used in a large sense
   Geometrical ratio of increase
   Nature of the checks to increase
   Complex relations of all animals and plants to each other in the struggle for existence
   Struggle for life most severe between individuals and varieties of the same species
chapter iv. natural selection; or the survival of the fittest
   Natural Selection
   Sexual Selection
   Illustrations of the action of Natural Selection, or the survival of the fittest
   On the Intercrossing of Individuals
   Circumstances favourable for the production of new forms through Natural Selection
   Extinction caused by Natural Selection
   Divergence of Character
   The Probable Effects of the Action of Natural Selection through Divergence of Character and Extinction, on the Descendants of a Common Ancestor
   On the degree to which Organisation tends to advance
   Convergence of character
   Summary
chapter v. laws of variation
   Effects of changed conditions
   Effects of the increased use and disuse of parts, as controlled by Natural Selection
   Acclimatisation
   Correlated variation
   Compensation and economy of growth
   Multiple, rudimentary, and lowly organised structures are variable
   A part developed in any species in an extraordinary degree or manner, in comparison with the same part in allied species, tends to be highly variable
   Specific characters more variable than generic characters
   Secondary sexual characters variable
   Distinct species present analogous variations, so that a variety of one species often assumes a character proper to an allied species, or reverts to some of the characters of an early progenitor
   Summary
chapter vi. difficulties of the theory
   Difficulties of the theory of descent with modification
   Absence or rarity of transitional varieties
   On the origin and transition of organic beings with peculiar habits and structure
   Organs of extreme perfection and complication
   Modes of transition
   Special difficulties of the theory of Natural Selection
   Organs of little apparent importance, as affected by Natural Selection
   Utilitarian doctrine, how far true: Beauty, how acquired
   Summary
chapter vii
   Miscellaneous Objections to the Theory of Natural Selection
chapter viii. instinct
   Instincts comparable with habits, but different in their origin
   Inherited changes of habit or instinct in domesticated animals
   Special instincts; Instincts of the cuckoo
   Slave-making instinct
   Cell-making instinct of the hive-bee
   Objections to the theory of natural selection as applied to instincts: neuter and sterile insects
   Summary
chapter ix. hybridism
   Distinction between the sterility of first crosses and of hybrids
   Degrees of sterility
   Laws governing the sterility of first crosses and of hybrids
   Origin and causes of the sterility of first crosses and of hybrids
   Reciprocal dimorphism and trimorphism
   Fertility of varieties when crossed and of their mongrel offspring not universal
   Hybrids and mongrels compared independently of their fertility
   Summary of Chapter
chapter x. on the imperfection of the geological record
   On the absence of intermediate varieties at the present day
   On the lapse of time, as inferred from the rate of denudation and of deposition
   On the poorness of our palaeontological collections
   On the absence of numerous intermediate varieties in any single formation
   On the sudden appearance of whole groups of allied species
   On the sudden appearance of groups of allied species in the lowest known fossiliferous strata
chapter xi. on the geological succession of organic beings
   On the slow and successive appearance of new species
   On extinction
   On the forms of life changing almost simultaneously throughout the world
   On the affinities of extinct species to each other and to living species
   On the state of development of ancient compared with living forms
   On the succession of the same types within the same areas, during the later Tertiary Periods.
   Summary of preceding and present chapter
chapter xii. geographical distribution
   Present distribution cannot be accounted for by differences in physical conditions
   Single centres of supposed creation
   Means of dispersal
   Dispersal during the Glacial period
   Alternate Glacial periods in the north and south
chapter xiii. geographical distribution -- continued
   Distribution of fresh-water productions
   On the inhabitants of oceanic islands
   Absence of Batrachians and terrestrial Mammals on oceanic islands
   On the relation of the inhabitants of islands to those of the nearest mainland
   Summary of the last and present chapter
chapter xiv. mutual affinities of organic beings: morphology -- embryology -- rudimentary organs
   Classification
   Analogical resemblances
   On the nature of the affinities connecting organic beings
   Morphology
   Development and embryology
   Rudimentary, atrophied, and aborted organs
   Summary
chapter xv
   Recapitulation and Conclusion
Glossary of Scientific Terms